The Titan Bestial, or Beast Titan, stands among the Nine Titans with a primal, ape-like visage in its current incarnation. Its physical traits adapt to each inheritor, though Zeke Yeager’s form looms larger than average Titans and wields heightened tactical intellect. This Titan exhibits lethal precision in hurling projectiles—boulders, debris—to devastate targets, alongside limited nape hardening and the power to transmute Subjects of Ymir into Pure Titans. By injecting spinal fluid and issuing vocal triggers, Zeke, leveraging his royal bloodline, commands Titan armies for warfare, exemplified during Wall Rose’s invasion and Fort Slava’s siege. Zeke inherited the Titan Bestial from Tom Ksaver under Marley’s Warrior program. Descended from the Fritz lineage, his royal blood grants authority over Pure Titans, directing their movements or suppressing their cannibalistic urges—a control less absolute than the Founding Titan’s. Titans under his sway function tirelessly under moonlight but sometimes defy orders, as seen when one ignored commands and consumed Mike Zacharias. Infiltrating Wall Rose during the Clash of the Titans arc, Zeke transformed Ragako villagers into Titans to probe Paradis’ defenses. At Utgard Castle, he shattered Wall fragments into lethal projectiles to decimate Survey Corps soldiers. His strategic mastery peaked at Shiganshina, where summoned Titans blocked retreats, and he catapulted Bertolt Hoover to unleash the Colossal Titan. Levi Ackerman’s ambush nearly ended Zeke, forcing a retreat via Cart Titan intervention. Zeke later deployed airborne Titanized Eldians in the Mid-East War, overwhelming enemy lines. Residual spinal fluid from his experiments altered Falco Grice’s Jaw Titan, imbuing avian traits. His ultimate aim centered on wielding the Founding Titan to sterilize Eldians, a euthanasia plan born from rejecting both Marleyan oppression and Eldian restorationist ideologies. The Titan Bestial’s saga culminates in the Rumbling arc. Zeke’s royal blood links him to Eren Yeager, activating the Founding Titan. Yet their ideological rift proves fatal: Levi severs Zeke’s spine, terminating his Titan command. Before death, Zeke manipulates memories within the Paths, bridging allies to thwart Eren’s annihilation campaign—a final gambit in a lifetime of calculated, conflicted loyalties.