Lolo is a young Adélie penguin born in Antarctica. His father, Toto, saved his egg during a skua attack that destroyed their other egg, making Lolo an only child. This fostered his inquisitive nature and persistent questioning, traits that sometimes frustrate older penguins like his grandfather, Pigo. Driven by curiosity, Lolo often explores beyond the safety of the penguin colony, usually alongside his best friend, Pepe, a pink penguin also raised as an only child.

His adventurous and fearless personality frequently leads him into danger. Early on, he wandered off to investigate his surroundings, such as attempting to befriend a baby skua—an action that provoked an attack from an adult skua, leaving him injured. During his recovery, he befriended a husky puppy named Don and encountered a scientist studying Antarctica, challenging the colony's distrust of humans. Despite paternal discipline for disobedience, including physical punishment, Lolo's exploratory behavior continued.

His development intensified when he and Pepe became stranded on an iceberg after an ice-breaking incident. This ordeal forced Lolo to confront predators like leopard seals and killer whales; he demonstrated quick thinking for survival, incapacitating a leopard seal with a snowball. Their subsequent capture by poachers tested his resilience further. Imprisoned aboard a ship with Pepe and a macaroni penguin named Mak, Lolo devised an escape, using a stick to open their cage latch. With help from the ship's St. Bernard dog, Jack, they fled during a storm and returned home via blue whales.

These trials culminated in Lolo's transition to leadership. After poachers attacked the colony to abduct penguin chicks, Lolo organized a rescue mission. Tragically, his father, Toto, died fighting the poachers during this effort, alongside other penguins killed by gunfire. In the aftermath, Pigo identified Lolo as the colony's new leader, a role endorsed by Lala, Pepe, and Mak. As winter approached, Lolo led the colony's migration to the North Sea, signaling his maturation from a curious chick to a responsible guide.

Throughout his journey, Lolo maintained key relationships: his steadfast bond with Pepe, his respect for Pigo's wisdom, his complex dynamic with his protective but stern parents, and his interspecies connections with Don, the scientist, and the whales that aided his return. His experiences with human threats—both benevolent researchers and violent poachers—shaped his understanding of the world beyond penguin society.