The Moguru soldiers descend from an ancient alien civilization that escaped their dying homeworld—a moon of Mars—seeking refuge in Earth’s oceans 200 million years before the central conflict. Governed by a rigid hierarchy, their society falls under the command of rulers like Hellqueen V and her successor, Queen Sandra, both orchestrating Earth’s conquest through calculated invasions. Among their ranks, the scientist Zanga devises colossal robotic monsters, while the steadfast Gorosky executes military strategies to devastate surface civilizations.

Militarily, the Moguru leverage subterranean strongholds, advanced technology, and genetically engineered mechanized forces, including an ancient army resurrected by Golaius, a legendary figure from their past. Their plans unravel when satellite surveillance exposes their underwater headquarters, triggering a decisive assault by adversaries. Central to their motives is the exploitation of "power Ypsilon," a rare genetic trait in humans that promises strategic supremacy.

The Moguru’s campaign culminates in a catastrophic final battle within their collapsing base, where Queen Sandra, Zanga, and key leaders meet their end, extinguishing their invasion ambitions permanently.