**Megatron** originated as D-16, a miner in Kaon’s depths who rose to fame as the gladiator Megatronus, later adopting the name Megatron. Initially rallying against Cybertron’s caste system, he established a tyrannical regime as Decepticon leader, shifting his ideology toward universal domination, believing Transformers superior to all weaker species. When the Unicron Singularity endangered Cybertron, he sought the Cyber Planet Keys and Omega Lock for absolute power instead of salvation, journeying to Velocitron, Jungle Planet, Earth, and Gigantion. He manipulated planetary rulers like Scourge and Override, seized Jungle Planet’s key, endured betrayal and usurpation by Starscream, then reclaimed command. On Gigantion, he harnessed the world’s key to become Galvatron, gaining enhanced strength and a new alternate form. Briefly allying with Planet X exiles Sideways and Soundwave for vengeance, he ultimately banished them to another dimension. Galvatron’s endgame involved using the Omega Lock to hasten the black hole’s collapse and reshape reality under his rule. After surviving a climactic battle within the black hole against united planetary leaders, he faced Optimus Prime alone in final combat and met his destruction.
**Red Alert** began as a fierce warrior in Cybertron’s civil war until a crippling injury cost him his hands, driving him to abandon combat for medical science. Emotionally detached but devoted to healing allies over fighting, he became the Autobots’ chief medic and scientist, partnered with Mini-Con Longarm. During the Unicron Singularity crisis, he relocated to Earth, adopting an emergency vehicle alternate mode, and served as Combat Medic in the Cyber Planet Keys quest. His caution frequently translated to tactical hesitation. After a near-fatal clash with Megatron, he was rebuilt with Hot Shot and Scattorshot into the Cybertron Defense Team, gaining upgraded armor and firepower while retaining medical capabilities. He aided in securing keys across multiple worlds and defended Cybertron during Primus’s awakening. Following Galvatron’s defeat and the black hole’s sealing, he continued serving the Autobots under Optimus Prime’s command.