Lily is a mouse Beastfallen with white hair and gray eyes, habitually wearing a hood that obscures her eyes. Her timid and shy nature drives her to avoid crowds and maintain a reserved presence. Though possessing a childlike appearance and a squeaky voice, she has lived over two decades, her aging decelerated by her Beastfallen heritage granting extended longevity.

She serves as a helper at a church, performing her duties with cheerful dedication fueled by loyalty and affection for the priest. This dedication reflects her inherently supportive character. Her mouse physiology grants heightened senses, including exceptional smell and hearing. She also possesses the unique ability to mentally commune with rodents, enabling her to direct them from a distance.

As a chef-in-training, she demonstrates remarkable culinary skill, crafting original recipes that rival experienced cooks. Beyond the kitchen, she exhibits proficiency in hand-to-hand combat and martial arts, effectively wielding a knife as her chosen weapon—a physical strength contrasting her usual timid demeanor.

Her Beastfallen identity causes deep self-consciousness, particularly regarding others' perceptions of her species. When asked to prepare meals publicly, she hesitates, expressing concern that her rodent nature might provoke disgust, revealing internalized anxiety about prejudice; reactions around her vary.

During a conflict involving church forces, Beastfallen mercenaries capture her, though she is later rescued, underscoring her vulnerability in danger. Her role within the church community remains steadfast, centered on support and service.