Millie Myers works alongside Yor Forger at Berlint City Hall. She possesses light skin, downward-slanting eyes accentuated by long lashes, and chin-length strawberry blonde hair styled in an inward-curling bob with a fringe swept right. Her hair shifts between blonde and orange across depictions, while her eyes vary between green and light blue. Millie wears the standard city hall uniform: a green vest over a white long-sleeved shirt, paired with a green skirt shorter than her colleagues', thigh-high black socks, and black heels. She accessorizes with a small red ribbon on her head's right side and round earrings. For social occasions like Camilla's party, she dons a strapless red dress featuring frills and a belt below the bust, a pearl necklace, red high heels with flower decals, and a thinner head ribbon.
Her personality is lively and mischievous, frequently smiling or giggling. She suggests pranks like putting snot in their section chief's coffee and maintains cheerfulness even after a breakup, immediately asking Yor to introduce her to the attractive Yuri Briar. Beneath this exterior lies deep emotional pain from her father's death during the second East-West war when she was very young. His absence left profound loneliness, which she blames for her difficulties and anger. Camilla notes Millie's dislike of being alone. This trauma caused her to avoid the previous year's charity bazaar for disabled veterans, as it reminded her of her loss.
Millie initially shares a strained relationship with Yor, joining Camilla and Sharon in gossiping about and insulting her. After Yor's marriage to Loid is revealed, Millie and the others engage Yor more, often offering dubious marital advice. Their dynamic shifts during collaborative events: when Millie and Camilla work overtime, Yor and Sharon assist, leading to their first group dinner. Millie witnesses Yor's drunken antics and heartfelt devotion to Loid, finding her behavior amusing and endearing. At the charity bazaar, Millie confronts Lady Patriots Society members criticizing her grief over her father, declaring soldiers survived while her father did not return. Yor defends her, acknowledging their generation's unique wartime suffering, deeply touching Millie.
Millie's resentment resurfaces upon learning of Yor's friendship with Melinda Desmond, wife of Donovan Desmond, whom she wrongly believes started the war that killed her father. She publicly accuses Melinda, prompting Matthew McMahon's correction that Donovan became prime minister after the war began. Humbled, Millie apologizes to Melinda, who empathizes with her pain. Millie later thanks Yor for her support, re-evaluating their relationship and changing her view of Yor from a "weird, clueless robot" to a "kindly gorilla." She chooses to continue helping at the charity event, valuing her co-workers' companionship over solitude.
Her background includes sharing her grief about her father with Camilla and Sharon. She dated a boyfriend who appreciated her short skirts, though they broke up prior to Jim Hayward's arrest at city hall. At Camilla's party, she mocked Yor for arriving without a partner but was stunned when Loid appeared as Yor's husband. She later expressed interest in Yor's brother Yuri, though unsettled by his sister complex. Millie's development culminates in acknowledging her reliance on her co-workers and actively choosing to participate in emotionally challenging events.