Yamato Mikoto hails from the Far East, orphaned in childhood and raised in an orphanage overseen by the deities Tsukuyomi and Takemikazuchi. She joined the Takemikazuchi Familia, where she and other members frequently smuggled Sanjouno Haruhime out of her manor to play, fostering a lasting friendship despite her father's disapproval that severed the shrine's relations. Years later, Mikoto relocated to Orario with the Familia to raise funds for the shrine, achieving Level 2 before the main events.
Mikoto possesses long black hair, usually in a ponytail, bluish-purple eyes, and wears a purple kimono with a sarashi underneath. Her attire features a dark red sash, thigh-length boots, a red shoulder guard on her non-weapon arm, arm guards, and a katana at her waist.
Her personality blends seriousness, honor, and humility. She spurns underhanded tactics, like disapproving of "Pass Parading" monsters onto allies. Despite her combat skill, she deflects praise and becomes embarrassed by it. Deeply loyal and caring, she prioritizes protecting loved ones, even employing stealth-based ninjutsu against her direct nature when required. She cherishes hot springs and adheres to specific bathing rituals.
As a Level 2 adventurer, Mikoto boasts enhanced strength, endurance, agility, and senses surpassing ordinary humans. Trained by Takemikazuchi, she masters multiple martial arts styles and weaponry, including katanas, spears, bows, axes, and thrown weapons like shuriken. Her signature techniques include "Mikazuchi," a modified aerial throw, and "Zekka," an advanced quick-draw strike capable of wounding formidable foes like the Monster Rex Amphisbaena. She practices Concurrent Chanting, enabling magic use while moving.
Her magic, "Futsunomitama," wields gravity manipulation to create a crushing dome-shaped field, effective against superior opponents, such as briefly immobilizing the Level 5 Black Goliath. Her Skills are "Yatanokurogarasu" for detecting previously encountered monsters and "Yatanoshirogarasu" for locating Familia members. She possesses the Development Ability "Abnormal Resistance" to negate poisons and similar effects. Her equipment includes katanas like "Kotetsu" and "Shunsan," forged by Welf Crozzo, and the short sword "Chizan," one of a pair gifted by Takemikazuchi.
Mikoto's relationships shape key arcs. Her lifelong bond with Haruhime compels her to join Bell Cranel in rescuing Haruhime from the Ishtar Familia, risking self-destructive magic use. She initially revered Takemikazuchi as a father figure, but her feelings deepened into romantic love. His gift of "Chizan"—promising its twin upon her Familia return—symbolizes their connection, with Mikoto intending to confess upon reunion. She maintains close ties with childhood friends from the orphanage, Kashima Ouka and Hitachi Chigusa, cooperating seamlessly in combat despite her temporary transfer to the Hestia Familia.
After her Familia's "Pass Parade" endangered Bell's party, Mikoto converted to the Hestia Familia to atone and support the War Game against Apollo. Though intended as a one-year arrangement, she gained respect for Bell and aided his efforts, including sheltering the Xenos. Her rapport with Hestia evolved from initial hostility to mutual respect, with Hestia praising her cooking and supporting her romantic pursuits.
Her character development centers on reconciling loyalties. While dedicated to Hestia Familia's endeavors—protecting Haruhime and aiding Bell—she plans to return to the Takemikazuchi Familia to fulfill her promise. Discussions note the practical value of her skills to Hestia Familia but acknowledge her deeper bonds and unresolved feelings for Takemikazuchi.