Usopp serves as the sniper of the Straw Hat Pirates and a Senior Officer of the Straw Hat Grand Fleet. Born in Syrup Village to Yasopp, a member of Shanks' crew, and the late Banchina, Usopp coped with abandonment and loss by telling elaborate lies and founding the "Usopp Pirates" with village children. His false stories entertained his sick mother and later his bedridden friend Kaya, though some contained elements of truth. This established his reputation as a chronic liar, undermining his credibility during genuine threats like Kuro and the Black Cat Pirates. The Straw Hats' intervention in Syrup Village led to his recruitment, with Kaya gifting the crew the Going Merry. Usopp dreams of becoming a "brave warrior of the sea" like his father, a goal challenged by deep-seated cowardice and insecurity about his usefulness compared to stronger crewmates.
Early adventures highlighted his technical ingenuity and sniping skills, but were often marred by fear-driven retreats and excuses like claiming "Can't-get-on-this-island disease." A significant turning point occurred during the Water 7 Arc. His emotional attachment to the damaged Going Merry and feelings of inadequacy culminated in a duel with Luffy and temporary departure. He returned under the alter ego "Sogeking" during the Enies Lobby Arc, symbolizing a psychological mechanism to access courage while maintaining distance. This period solidified his role as a critical support fighter using slingshots and improvised gadgets. Meeting the giants Dorry and Brogy in Little Garden ignited his desire to visit Elbaph and learn from its warrior culture.
The two-year timeskip marked substantial growth. Training on the Boin Archipelago transformed him physically—increasing muscle mass, adding a goatee, and lengthening his hair—and enhanced his arsenal with Pop Greens (combative plants) and dial-based ammunition. While retaining his trademark cowardice and susceptibility to scams, he displayed newfound combat confidence, such as attacking the Fake Straw Hat Crew at Sabaody without hesitation. His sniping goggles were replaced by headphones and a white sun cap, and his yellow satchel became red. Despite declaring himself graduated from the "Weakling Trio," he continued to panic in high-risk situations, opposing confrontations with entities like the Four Emperors and pleading for lower bounties to avoid stronger enemies.
In the Fish-Man Island Saga, Usopp warned the crew about the Kraken during the descent from Sabaody and later devised strategies to fight underwater without damaging the ship's protective coating. On Fish-Man Island, he battled Daruma, an officer of the New Fish-Man Pirates, using Pop Greens like the "Sasuke Matryoshika" to immobilize opponents. He remarked on his diminished reliance on bluffs, noting his abilities now matched his boasts. He also exhibited mild perversion alongside Zoro and Brook when trapped in Ryugu Palace, reacting to Brook's soul-based surveillance. His loyalty surfaced when he protected Shirahoshi and King Neptune.
Post-Fish-Man Island, his bounty rose to 200,000,000 berries after the Dressrosa Arc—where he adopted the alias "God Usopp" and inspired dwarves as a descendant of Noland—and later to 500,000,000 berries after the Raid on Onigashima. Despite this recognition, he remained conflicted, viewing higher bounties as dangerous liabilities. His cowardice persisted, such as nearly abandoning allies during Operation S.O.P. in Dressrosa, though he ultimately resisted. Conversely, he demonstrated fierce resolve in Wano by refusing to abandon injured allies like Kin'emon and Kiku, berating their acceptance of death for honor and emphasizing survival. His willpower allowed him to withstand Big Mom's Conqueror's Haki release.