Daisuke Jigen pursues the legendary Columbus Egg—a treasure rumored to wield weather control and immense value. This quest ignites after antagonist Nazaroff attacks Fujiko Mine, who memorized the Columbus Files' location details. With the files destroyed and Fujiko stricken by memory loss, she becomes the sole key to the egg. Jigen allies with Lupin III and Goemon Ishikawa XIII to recover the treasure and restore Fujiko’s memories.
His role leverages firearms expertise during the operation, though specific combat actions remain undefined. He engages in characteristic banter with Goemon, echoing their established dynamic. The narrative spotlights the team’s collective efforts to surmount obstacles in the treasure hunt and Fujiko’s plight.
Clad in his trademark dark suit and fedora, consistent with his franchise appearance, Jigen bolsters the team’s navigation of diverse locales and confrontations against Nazaroff’s forces. Their dual objectives drive them: securing the Columbus Egg and aiding Fujiko.