Yelena, a tall androgynous figure with a sharp blonde bob and a faux chinstrap beard, transitioned from a Marleyan military uniform to crisp civilian attire—a white button-down paired with a dark blazer and slacks. Born into an unassuming Marleyan family, her disillusionment with the nation’s imperial cruelty drove her to craft a false history of her homeland’s subjugation by Marley, weaponizing this narrative to fuel her insurgency. Her destiny pivoted during the Marley Mid-East War when Zeke Yeager rescued her unit from annihilation, igniting an obsessive loyalty to him. She embraced Zeke’s euthanasia scheme—sterilizing Subjects of Ymir to eradicate Titan inheritance—as divine prophecy, dedicating herself as his most fervent disciple. Commanding the Anti-Marleyan Volunteers, Yelena brokered strategic pacts with Paradis Island, supplying arms and tactical expertise to destabilize Marley. She spearheaded the Liberio Raid, infiltrating enemy ranks disguised as a soldier to incapacitate Pieck and Galliard, securing Eren’s assault. Her covert operations included tainting Paradis officials’ wine with Zeke’s spinal fluid, granting him remote dominion over their bodies. While posing as a loyal ally to Paradis, she privately manipulated Eren, steering him toward Zeke’s agenda while obscuring her machinations from military leaders. Her fanaticism spurred ruthless acts: executing comrade Griez for mocking Eldians and orchestrating Dot Pixis’ political ruin after his dissent. Even after Zeke’s death and Eren’s betrayal, she clung to the euthanasia ideal as the Rumbling raged. Captured by Floch’s faction and condemned for her role in Zeke’s plots, she was liberated by the Cart Titan, joining the coalition against Eren and revealing crucial intel to cripple Fort Salta’s air defenses. Post-conflict, Yelena endured, her zeal muted by devastation. She endeavored to propagate Zeke’s vision despite its collapse, her crusade reduced to whispers of a failed utopia. Ultimately, she fled aboard a lifeboat with surviving remnants of the old world, her fire dimmed but unextinguished.