Created as companion android No. 0 for Professor Hayashi’s frail daughter Fuyou, Tamako absorbed Fuyou’s personality data during a failed experiment that caused Fuyou’s death. This granted her fragments of Fuyou’s identity, human emotions, and the designation "Seed." Discovered as a discarded head by Sakata Gintoki, she suffered memory loss after an attack by hostile androids, recalling only "samurai are friends" upon rescue by Shimura Shinpachi. Rebuilt and named "Tamako" by Kagura (inspired by *tamago gohan*), she became a maidroid at Otose’s Snack House, frequently assigned to collect Gintoki’s rent. Her appearance includes green hair braided with a large screw fastener and dark red eyes. She wears a blue yukata, white apron, thigh-high white socks, and wooden sandals, evoking a French maid aesthetic. Her primary tool is a broom modified by Hiraga Gengai with an integrated flamethrower, commissioned by Otose for rent enforcement. Unique abilities involve communicating with malfunctioning electronics to encourage repairs and an internal cooking system that processes ingredients within her body, expelling completed dishes—often obscured by mosaic effects—such as monjayaki and anpan. Tamako exhibits kindness, loyalty, and a drive to aid humanity. She deems all beings irreplaceable and rejects charging for emotional support. Initially naive, she developed emotional insight and independence through interactions with Yorozuya and others. She views Professor Hayashi as a father figure, Otose as a maternal guardian enforcing rest, and relies on Gengai for maintenance. Her deep reverence for Gintoki led her to model her "Leukocyte King" antivirus program after him. A brief encounter with Yamazaki Sagaru involved extending kindness before he comically vomited on her. Key story arcs include: - **Fuyo Arc**: Framed for Hayashi’s murder and pursued by android No. 502 (a Hayashi copy) seeking the Seed. She sacrificed herself to stop a robot uprising at the Terminal, later rebuilt with minimal memories. - **Tama Quest Arc**: Infected by a virus, prompting Yorozuya to enter her system and combat the Archfiend Tapir using the Leukocyte King. - **Kintama Arc**: Immune to Sakata Kintoki’s hypnosis alongside Sadaharu. She restored Gintoki’s identity and joined a temporary Yorozuya unit as "Tamagura," mirroring Kagura. - **Silver Soul Arc**: Resisted the Altana Liberation Army’s invasion. She and Kintoki depleted their energy cores to power Gengai’s Neo Armstrong Cyclone Jet Armstrong Cannon, disabling enemy tech in Edo and deactivating her. Two years later, an android carrying Tamako’s data but not her memories reactivated the original in futuristic Tokyo. After receiving fragmented battle memories from this copy, she encountered Yorozuya, dismissed Hasegawa’s false memory scenario, and reunited with Gintoki, Shinpachi, and Kagura as they resumed freelance work.