The Misaka Sisters are genetic clones of Mikoto Misaka, originating from the clandestine Radio Noise Project. Initially designed as Level 5 esper replicas, their purpose pivoted to Accelerator’s Level 6 Shift experiment, requiring his elimination of 20,000 clones to ascend. Identical to Mikoto in form, they bear hollow gazes, cropped hair, and signature gear: electromagnetic-perception goggles and blue-striped undergarments. Artificial cultivation tanks and hormone treatments accelerated their growth, shortening lifespans until post-experiment medical interventions stabilized them. Programmed with emotional inhibitors during the experiment, they communicated in monotone third-person speech, displaying muted affect. Kamijou Touma’s disruption of the project—including rescuing Misaka 10032 from Accelerator—sparked a transformation, unlocking latent emotional agency. Subsequent clones cultivated individuality, with some adopting human-like behaviors such as dieting or crafting gifts to court Touma’s favor. Their interconnected consciousness, the Misaka Network, operates as a real-time hive mind, pooling memories and senses. Beyond collective awareness, it serves as a dynamic processing unit—refining Mikoto’s railgun trajectories or offsetting Accelerator’s cognitive deficits. Last Order (Misaka 20001), the network’s administrator, safeguards against data corruption. Rarely, the network coalesces into a unified sentience called the "Will of the Misaka Network," demonstrating nuanced emotions and independent decision-making. Clones wield Level 2-3 electromaster powers, excelling in electronic disruption, infiltration, and projectile manipulation. Testament machines hardwired combat protocols into their neural frameworks, granting expertise with firearms like the F2000R Toy Soldier and Metal Eater MX. Though weapon mastery propagates network-wide, distinct personalities emerge: Misaka 10032 nurtured affection for a stray cat and a pendant from Touma, while Misaka 10046 grappled with fear and shifting allegiance toward Accelerator. Post-experiment survivors scattered globally for specialized care, though some remain in Academy City. Freed from systemic control, they explore hobbies, diet competitions, and social dynamics. Interactions with Last Order and Accelerator oscillate between rivalry and guardianship, reflecting evolving relational complexities. The network’s utility extends to crisis stabilization and occasional communion with metaphysical entities like artificial angels, underscoring its adaptive significance beyond computational roles.