Professor Sycamore, called Professeur Platane in Japanese-language media, holds the position of the Kalos region's Pokémon Professor in Lumiose City. His central research pursuit is Mega Evolution, a phenomenon he studies through both field observations and laboratory analysis. Trained under Professor Rowan in Sinnoh, he later moved to Kalos and studied at the Tower of Mastery to comprehend Mega Evolution. Ultimately concluding he lacked the personal strength to wield it, he dedicated himself to scholarly research. He exhibits a consistently calm and supportive demeanor, prioritizing Pokémon well-being and aiding trainers in understanding their partners. This is shown when he patiently nurtured an injured Gible that bit him, guiding it to evolve into Garchomp. His commitment to fostering trainer-Pokémon bonds extends to mentoring assistants and new trainers, demonstrated by his gifts of starter Pokémon—a Charmander to Alain and Kalos starters to Tierno, Shauna, and Trevor. His relationship with Alain is significant; he supported Alain's quest to collect Mega Stones despite the latter's abrupt departure from the lab, maintaining periodic communication and showing concern for his progress. This dynamic shifts in the Mega Evolution Specials, where Professor Sycamore expresses visible shock learning of Alain's battles against Primal Groudon, Primal Kyogre, and Mega Rayquaza in Hoenn. He later observes Alain's participation in the Lumiose Conference, noting his cold demeanor and attempting to reconnect afterward. Interactions with Lysandre de Fleur-de-lis reveal a layer of naiveté; initially trusting Lysandre as a colleague researching Mega Evolution's peaceful applications, he is stunned to discover Lysandre's leadership of Team Flare and manipulation of Alain. During the Team Flare crisis, he actively participates in counter-operations, rescuing Mairin from grunts and collaborating with allies like Steven Stone, Diantha, and Gym Leaders to storm Prism Tower. In *The Strongest Mega Evolution ~Act IV~*, he appears supportively, discussing Alain's escalating battles and Lysandre's motives, displaying concern for Alain's obsessive pursuit of strength and the well-being of Manon's Chespin after it encounters a mysterious green entity at Lysandre Labs. This reinforces his role as an observer and commentator on Mega Evolution's broader implications. His primary Pokémon partner is a Garchomp, raised from the injured Gible. This Garchomp can Mega Evolve using a Garchompite and a Keystone worn on Sycamore's watch band. He utilizes Mega Garchomp in critical situations, such as countering Team Flare attacks or protecting allies.