Fleur-de-lis, alias Lysandre, commands Team Flare as its enigmatic leader and directs Lysandre Labs in the Kalos region. His mission centers on controlling Mega Evolution information to curb its misuse through calculated manipulation and covert operations. He first crosses paths with Alain during a search for a Denryunite in ancient ruins, offering the stone as a prize for victory in their battle. After defeating Alain, he enlists him under the pretense of safeguarding Professor Sycamore’s reputation, supplying a Key Stone and Charizardite X while concealing his deeper agenda. Later, Lysandre orchestrates Alain’s training through a grueling gauntlet of battles against ten Mega Evolution users, threatening revocation of his Mega Ring upon defeat. This trial serves dual aims: assessing Alain’s determination and harvesting data on Mega Evolution’s energy patterns. Simultaneously, he partners with scientists to analyze the Giant Rock, a relic emitting ancient energy signatures, which he isolates in a containment unit to avert regional disaster. Exchanges with Steven Stone expose his methodical curiosity about historical artifacts and Mega Evolution’s roots, yet he skillfully sidesteps inquiries into his ultimate objectives. Experiments involving Zygarde Cores and green cellular entities spiral into chaos when Chespie, a Chespin in his custody, falls comatose after absorbing volatile energy. While publicly vowing to rehabilitate Chespie, Lysandre covertly leverages the crisis to coerce Alain into accelerating research, framing the completion of the Mega Evolution system as Chespie’s sole hope—a tactic highlighting his subordination of ethics to scientific progress. Tensions simmer with Kalos Elite Four member Malva, who probes his intentions during Alain’s climactic battle. Lysandre deflects her skepticism by redirecting focus to Alain’s triumphs. Privately, he masterminds Zygarde Core captures and tracks the Giant Rock’s energy fluctuations, methodically advancing his vision to restructure society through calibrated power application. Lysandre’s battle strategy mirrors his calculated nature, deploying Pyroar and a Shiny Mega Gyarados armed with Hyper Beam and Incinerate. These confrontations double as trials for allies like Alain and exhibitions of his martial prowess. His cross-media narrative consistently threads a philosophy of eradicating humanity’s perceived imperfections, though the anime iteration emphasizes subterfuge and data-driven machinations over the apocalyptic overtures of his game counterpart. Ancestral connections to Kalosian royalty surface in game lore, while anime canon leaves his history nebulous—no confirmed ties to Professor Sycamore precede their documented encounters. This duality as both patron and adversary weaves intricate dynamics, blending Alain’s mentorship with shadowy maneuvers that incrementally realize his ideological blueprint.