Fiorellino, first called Honey Honey, is an orphaned teenager working as a Vienna waitress. Abandoned as an infant in a meadow teeming with honeybees, nuns discovered her there, named her Honey, and raised her in their convent. This echoes her own discovery of a similarly abandoned white cat, Lily, whom she regards as a sister, not merely a pet.

She embodies kindness, resilience, and a powerful protective instinct, especially towards Lily. This devotion drives her to turn down lucrative offers for the cat, unaware Lily swallowed Princess Flora's priceless ring. She prioritizes Lily's safety above all else, enduring imprisonment and hunger strikes when separated.

Her journey becomes a global pursuit after Lily ingests the ring. She evades Princess Flora's suitors and the gentleman thief Phoenix across continents. During this ordeal, romantic feelings for Phoenix emerge, shown when she murmurs his name in her sleep. Captured by a nomadic tribe, they force her to walk barefoot over hot coals, revealing a rose-shaped birthmark on her foot. This confirms her true identity as Princess Priscilla, younger sister of Princess Flora and rightful heir to a small Prussian kingdom destroyed years earlier by the antagonist Slag.

Following this revelation, Slag imprisons her in a Siberian castle. She escapes with Phoenix's aid and reunites with her father, now a Moscow gardener. Their subsequent journey leads to enslavement in Constantinople. Sold to an Indian sultan fascinated by magic, she seizes a chance to steal his magic carpet, enabling further travels. The climax occurs in New York, where she reconciles with Flora after their sisterhood is acknowledged. Demonstrating her compassion, she peacefully resolves King Kong's capture of Flora. Her story concludes with marriage to Phoenix, revealed as a nomadic tribe leader's son, and an understanding of the prolonged pursuit's significance concerning Lily.