Apos stands as the primary antagonist, an agender immortal angel commanding fellow angels to persecute immortals. As Yggdrasil’s current guardian—the mythical tree bearing Time Fruits—Apos plots to sacrifice Rin Asougi to achieve permanent guardianship. This scheme aims to consume Rin’s Time Fruit and absorb her centuries of accumulated memories.

Born to Tajimamori, Yggdrasil’s prior guardian, Apos inherited the role but defies the natural transition cycle, seeking eternal power through Rin’s sacrifice. Intersex by nature, Apos bears both male and female secondary sex characteristics, a trait believed to confer divinity. This fuels a godlike self-perception devoid of human morality, manifesting in extreme sadism. Apos tortures immortals trapped in Yggdrasil’s roots, convinced suffering enriches their Time Fruits’ flavor.

Over decades, Apos hunts Rin using diverse methods, including mortal assassins like Laura. Uniquely, Apos can instantly extract Time Fruits from immortals or angels, inflicting true death—a power others lack, signifying profound ties to Yggdrasil’s mechanics.

In the final clash, Rin overpowers Apos, forces Tajimamori’s Time Fruit into them, and hurls them into Yggdrasil’s core. This severs Apos’s guardianship and triggers permanent imprisonment. Facing absorption, Apos unravels into desperate pleading. Now eternally trapped within the tree’s consciousness, Apos suffers without hope of rescue.