Oswald Cobblepot, the Penguin, is displaced from modern Gotham City to feudal Japan by a temporal anomaly. He ascends as a warlord, securing territory and forces while constructing a unique mecha castle. Its glacier-like interior reflects his avian motif, offering familiarity in this era.

He joins a coalition of Gotham villains ruling partitioned regions under the Joker's supreme command. This alliance coordinates dominance through territorial warfare, including clashes with adversaries like Poison Ivy, who controls adjacent lands.

During the final conflict, his mecha castle combines with others to form the colossal "Lord Joker" battle robot, designed to eradicate Batman and allies. This fusion embodies the villains' strategic consolidation under the Joker. The coalition falls to the Bat-Ninja clan and Gorilla Grodd's monkey ninja army. Defeated, he returns to the present with fellow villains and is apprehended by authorities.

He retains his aristocratic poise and strategic criminal methodology throughout, adapting operations to leverage feudal power structures and mecha technology. His role remains that of a calculated contributor to collective villainous ambitions, never the principal architect.