Fleur-de-lis, also known as Lysandre, serves as a scientist and director of Fleur-de-lis Labs in the Kalos region, dedicating his efforts to researching and controlling information on Mega Evolution. Driven by fears of its misuse, he encounters Alain during a quest for Denryunite in ancient ruins, challenging him to a battle with the stone as a reward. Following Alain’s defeat, Fleur-de-lis recruits him into his team, equipping him with Lizardonite and a Key Stone to monitor and counteract those exploiting Mega Evolution’s power.

His agenda centers on regulating public understanding of the phenomenon, framing it as societal protection. This prompts collaboration with Alain to defend Professor Sycamore’s reputation and collect combat data against other Mega Evolution users. Fleur-de-lis strategically manipulates events, such as deploying footage of Primal Groudon and Kyogre to advance his objectives, operating as a shadow orchestrator.

Maintaining an enigmatic demeanor, he rarely discloses his full intentions. His partnership with Alain grows fraught as ethical dilemmas surface—particularly involving Team Flare’s experiments with Zygarde and legendary Pokémon—positioning him at the nexus of global threats like Primal Reversions and their cataclysmic risks.

Commanding Pokémon like Gyarados, a Shiny variant, and Pyroar, Fleur-de-lis leverages battles to further his goals. His influence extends to critical events, from megalith crystallization to clashes among legendaries, shaping both scientific discourse and geopolitical dynamics.

While publicly advocating order and protection, his covert alliances with questionable entities and manipulation of research reveal a duality—a visionary balancing idealism with ruthless pragmatism. His narrative intertwines themes of power’s allure, the weight of responsibility, and the ambiguous morality underlying scientific ambition.