Haruka Haruno is the central protagonist of her season, a thirteen-year-old girl who enters the prestigious Noble Academy with a long-held and deeply personal dream. From a young age, she has been inspired by the princesses in her cherished picture book, viewing them as the embodiment of grace, kindness, and the power to make dreams come true. This admiration blossomed into her own greatest aspiration: to become a princess herself. However, this dream was not always met with support. A painful memory from her childhood, when a classmate mocked her ambition, made her initially shy about sharing her true feelings with others.

Despite this early discouragement, Haruka is defined by her cheerful, energetic, and remarkably persistent personality. She possesses a radiant smile and an unshakeable belief that to achieve one's dreams, hard work and perseverance are essential. She is fiercely determined, refusing to give up even when faced with seemingly insurmountable obstacles. While she can be clumsy and sometimes struggles academically or with her princess lessons, her enthusiasm is boundless, and she is always willing to try new things. This can occasionally lead to reckless or messy results, but her core nature is one of resilience; after moments of doubt or hurt, she always finds the strength to stand back up and continue pursuing her goals. Her characteristic phrases include exclaiming that something is "too amazing" and declaring that her feelings or a situation are "in full bloom".

Haruka's journey as a Cure Flora begins when she encounters two fairies, Pafu and Aroma, and witnesses the forces of Dyspear attempting to crush a classmate's dream. Her strong desire to protect that dream awakens her latent powers, allowing her to transform for the first time. Her role in the story is that of the leader of the Precure team, the Princess of Flowers, who fights to protect the dreams of all people from being sealed away in cages of despair. Her motivation is intrinsically tied to her personal belief system; she fights not only to save others but to uphold the very idea that dreams are precious and worth striving for, directly opposing the villains who seek to spread despair. As the story progresses, her understanding of what it means to be a princess evolves. She learns that true princesshood is not about status or a fairy-tale title, but about having an inner strength and nobility of character, and using that strength to inspire and protect others.

Her key relationships are central to her growth and the narrative. At school, she quickly befriends her roommate, Yui Nanase, whose dream of becoming a children's book author is the first one Haruka saves as a Precure. She forms a powerful bond with her teammates, Minami Kaido and Kirara Amanogawa. Initially, she is intimidated by the seemingly perfect Minami but grows to become her close and trusted friend. With Kirara, she is persistent and admiring, even helping her with work to convince her to join the team, and Kirara gives her the affectionate nickname "Haru Haru". A pivotal figure in her past is Prince Kanata from the Hope Kingdom, who met her as a child when she was crying after being mocked. He gave her a Dress Up Key and encouraged her to never give up on her dream, making him a lasting source of inspiration. Later, she plays a crucial role in saving Towa Akagi, who was under Dyspear's control. Haruka becomes a positive and uplifting presence in Towa's life, helping her find her own path as a Cure. Her family, including her supportive mother Moe, her father Ibuki who owns a Japanese confectionery shop, and her younger sister Momoka, also provide a foundation of love and encouragement, though her relationship with Momoka has its own storyline where they reconcile after a misunderstanding.

Over the course of the series, Haruka demonstrates significant development. She grows from a girl who was shy about her dream into a confident leader who proudly proclaims her ambitions. Her journey involves learning that the path to a dream is filled with challenges and setbacks, and that true strength comes from facing those challenges with the support of friends. By the epilogue, she has matured, having learned that her dream of being a princess is realized not through a title, but through her actions and her unwavering dedication to protecting the hopes of others.

As Cure Flora, her notable abilities are themed around flowers and elegance. She transforms using her Princess Perfume and various Dress Up Keys. Her main attack is Floral Tourbillon, typically performed after shifting into Mode Elegant, where she releases a torrent of flower petals to purify her enemies. With different keys, she can perform other powerful attacks such as Rose Tourbillon using the Crystal Princess Rod, and Sakura Turbulence. Her appearance as Cure Flora changes to reflect these different modes, with her hair growing long and turning blonde with pink gradients, and her dress becoming an elaborate, flowing gown.