Vector Prime is one of the Thirteen original Transformers created by Primus billions of years ago to combat Unicron. As the appointed guardian of time and space, he exists as a multiversal singularity, observing and recording events across countless realities while maintaining the timestream's integrity. His prolonged isolation from linear existence renders him culturally detached from modern Cybertronian society, often expressing himself in archaic, chivalric language, yet he remains fiercely dedicated to preserving all life.
He manipulates space and time for interdimensional travel, temporal observation, and localized time reversal. His primary tool is the sword Rhisling, forged from ethereal supermatter, capable of slicing reality to open dimensional or temporal portals. He employs a temporal mechanics analyzer to perceive multiple timelines, with internal inhibitors preventing paradoxes, though excessive power use risks physical dissolution. He transforms into an ancient Cybertronian spacecraft armed with gauss cannons and a causality-disrupting probability torpedo. His partners are the Mini-Con Safeguard, who mounts as a blaster, and the Recon Mini-Con Team.
In the *Cybertron* animated series, Vector Prime re-enters the timestream to counter the Unicron Singularity, allying with Optimus Prime's Autobots and providing knowledge of the Cyber Planet Keys. Initially deceived by Megatron, he surrenders a cosmic map to the Decepticons. Key actions include saving Landmine by warping him to Earth and identifying the Omega Lock symbol on a human child's clothing, guiding the artifact search. He aids in securing keys on Velocitron and the Jungle Planet. His ultimate sacrifice occurs on Gigantion, rewinding time to reopen a vortex, knowing it will fatally overextend him. Before dissolving, he transfers his planetary map to the human Bud and releases Safeguard. Later, in a non-corporeal state, he fortifies Optimus Prime within the singularity and bequeaths Rhisling to ensure Galvatron's defeat.
Fun Publications comics expand his role: he repairs timeline damage from Predacon Megatron during the Beast Wars and opposes Unicron's heralds like Ramjet and Nemesis Prime. After Unicron's destruction creates the singularity, he collaborates with Alpha Trion and Sentinel Maximus against herald-led assaults on Primus. His history includes briefly holding the Matrix of Leadership in an alternate reality and assisting Nexus Prime in reassembling the Star Saber, an act that splintered the Thirteen. The "Ask Vector Prime" feature details his multiversal insights, including an incarnation operating covertly on Nebulos with a Nebulan partner also named Safeguard.
In the Aligned Continuity (*Robots in Disguise*), Vector Prime resides in the Realm of the Primes, contributing spark energy with five other Primes to empower Optimus Prime against Megatronus. A separate incarnation appears in the *Revenge of the Fallen* film as one of the Primes betrayed by the Fallen.
Vector Prime’s legacy endures as a spectral guardian. In his final animated appearance, he manifests as an apparition to encourage Optimus Prime, with closing scenes depicting him eternally battling Galvatron’s spirit across time, symbolizing the perpetual struggle between order and chaos.