Override hails from Velocitron, the Speed Planet, where leadership is won through racing supremacy. As the planet's undisputed fastest racer, she claimed leadership, embodying Velocitronian culture with decisive action, self-assurance, and an unending drive for challenges. This initially surfaced as cockiness and occasional selfishness, focused solely on defending her racing title. Her path to power involved entering the leadership race disguised as male—a necessity under Velocitron's societal norms—revealing her true identity only after victory.
Interactions with the Autobots, especially Hot Shot and Optimus Prime, profoundly shaped her development. Races against Hot Shot and witnessing Autobot honor, such as Hot Shot sacrificing his lead to rescue her from a collapsed tunnel, transformed her from a fiercely competitive racer into a more compassionate and responsible leader. While her arrogance diminished and her focus shifted towards her people's welfare, she retained her competitive fire and love for speed. A close bond formed with the human girl Lori, whom Override calls "Little Sister," with Lori reciprocating as "Big Sister," revealing Override's nurturing side and strengthening her connection to Earth.
In the primary Cybertron storyline, Override initially withheld Velocitron's Cyber Planet Key, distrusting both Autobot and Decepticon intentions. She challenged Megatron and Hot Shot to races for information or the Key itself, demonstrating her speed and tactical acumen. Sabotage by Decepticon allies Ransack and Crumplezone during the Planet Cup race tested her resilience. Hot Shot's selfless actions during the event ultimately convinced her of Autobot integrity. After losing a fair finale race to Hot Shot, she relinquished the Key, joined the Autobots, and aided their mission by inserting Velocitron's Key into the Omega Lock, participating in subsequent quests to Jungle Planet and battles against Decepticon forces.
Across other continuities, her role expands. In IDW Publishing's 2005 universe, she co-leads Velocitron with Ransack amidst political tension. She approves Velocitron's entry into the Council of Worlds following racer Blurr's victory in the Benefit 500, affirming her commitment to galactic ties. Her story concludes tragically when Unicron devours Velocitron, her body depicted among the casualties. The 2019 IDW continuity shows her competing in the Speedia 500, losing to Mayhem, and later witnessing political upheaval at a ceremony. The prose story Last Bot Standing places her in a dystopian future, serving ex-Decepticon Steeljaw and participating in biofuel harvesting on Donnokt, where she encounters Rodimus and confronts moral ambiguities.
In Aligned continuity novels, Override rules Velocitron but contends with planetary decline, including a dying sun and civil unrest. She allies with Optimus Prime after the Ark's arrival, supporting the Autobots despite their refugee status. When civil war erupts, instigated by rival Ransack's Decepticon allegiance, she leads her faction against him, pledging loyalty to the Autobot cause before their departure. Beast Wars: Uprising reimagines her as a classist Cybertronian administrator overseeing gladiatorial games; her elitism sparks conflict with Maximal rebels.
A notable distinction exists in regional portrayals: the Hasbro/Cartoon Network version depicts Override as female, while the Japanese Galaxy Force iteration presents the character as male under the name Nitro Convoy. This difference influences certain dynamics, such as her relationship with Lori being sisterly in the former and formal in the latter.