Roni is a stern and uptight cleric who holds a deep disdain for demi-humans, making him an unlikely companion for the newest member of his adventuring party. Amona is a dark summoner, a demi-human who wields demonic powers and whose very existence seems to challenge everything Roni stands for. The rest of the party, including the elf archer Eury and the naive warrior Haru, brace for constant conflict between the priest and the summoner. However, to everyone’s astonishment, Roni and Amona end up spending their very first night together as a party. Following this unexpected encounter, the two begin a secret physical relationship while continuing to clash during their adventures, all while trying to hide their growing connection from their companions.

The story is set in a fantasy world that draws on classic Japanese role-playing game tropes, where adventuring parties take on quests, battle monsters, and navigate the politics of various races. Roni and Amona come from opposing backgrounds, yet their mutual attraction repeatedly draws them together. A significant source of comedy comes from their attempts to maintain a professional front, sneaking away from the group under thin excuses. Their secret is an open secret to the other party members, particularly Eury, who grows frustrated with their transparent behavior and the noise complaints their nighttime activities generate from innkeepers. Amona can summon an imp familiar, a small demonic creature that often serves as an unwitting commentator on her relationship with Roni.

A central narrative arc involves the gradual shift from a purely physical arrangement to genuine emotional attachment. Roni struggles with his religious upbringing, which taught him to reject earthly desires, while Amona deals with the prejudices faced by demi-humans. The party takes on various missions, including encounters with creatures like orcs that subvert typical fantasy expectations. As the volumes progress, the group dynamics evolve, and what begins as a one-night stand develops into a more complex relationship that the pair slowly stops hiding from their friends. The series combines romantic comedy elements with erotic situations, using the fantasy adventure setting as a backdrop for exploring the unlikely pairing of a cleric and a dark summoner.