Perona, known as "Ghost Princess," appears across multiple One Piece narratives, including the film *One Piece: Stampede*. Her background traces to commanding Wild Zombies and Surprise Zombies under Gecko Moria on Thriller Bark, where she served among the Mysterious Four and acted as a primary antagonist during the Thriller Bark Saga. Following Thriller Bark's collapse, Bartholomew Kuma's powers transported her to Kuraigana Island. She resided in Dracule Mihawk's castle, tending to Roronoa Zoro's injuries during their two-year stay and developing a cooperative yet contentious relationship with him. Perona exhibits a personality marked by immaturity, selfishness, and aversion to control or disobedience. She displays deep fondness for cute objects and creatures, aspiring to command a kingdom of adorable zombie animals. Despite a typically cruel demeanor, she demonstrates loyalty to Moria, viewing him as a father figure, and shows moments of kindness like nursing Zoro. Her abilities derive from the Horo Horo no Mi (Hollow-Hollow Fruit), a Paramecia-type Devil Fruit enabling her to generate ghosts with specific effects: Negative Hollows drain willpower, Mini Hollows explode on command, and Special Hollows cause larger destruction. She can project her spirit form, gaining intangibility and flight while leaving her physical body vulnerable. In *One Piece: Stampede*, Perona appears during the Pirates Festival on Delta Island alongside Dracule Mihawk and Boa Hancock amid chaos involving Marines and pirate factions. She employs her Negative Hollows against Marine forces during the large-scale battle, contributing to the pirate alliance's defense against the Buster Call fleet. Her design retains her post-timeskip appearance: a long black strapless dress with lavender frills, red high-heeled boots, a wide-brimmed black hat with floral accents, her signature parasol, and pink hair in spiral braids. She carries a stuffed replica of Kumashi, reflecting her affinity for cute aesthetics. Perona's broader journey includes departing Mihawk's castle upon learning of Moria's survival to reunite with him. Her interactions highlight themes of reluctant alliances and gradual maturity, while her core traits—childlike whims, katsaridaphobia (fear of cockroaches), and unique "Horo-horo-horo" laugh—remain consistent.