Mikoto Misaka, a 14-year-old Level 5 esper ranked third in Academy City, wields the electromaster ability Railgun. Daughter of Misaka Tabigake and Misaka Misuzu, she enrolled in elementary school, ascending from Level 1 to 5 through relentless training. Her DNA fuels the Sisters Project, a cloning initiative she endorsed under the deception of aiding muscular dystrophy research, spawning over 10,000 clones. Discovering their intended use in lethal experiments triggered profound guilt and trauma. Her electromastery enables hypersonic metal projectiles, lightning generation, and iron sand barriers. She manipulates electronics, senses electromagnetic fields, and disrupts telepathy by interfering with neural signals. A distinctive quirk is her habitual pairing of shorts beneath skirts, often remarked as unconventional. Initially prideful and competitive, her worldview fractured during the Sisters Arc. After confronting Accelerator, the top Level 5, to halt the clones’ slaughter, her defeat spiraled into a resolve for self-sacrifice—averted only by Kamijou Touma’s intervention. This ordeal reshaped her into a humbler figure reliant on allies and fiercely protective of the clones she now calls siblings. Her dynamic with Touma evolved from rivalry over his power-negating ability to a tangled bond marked by unspoken affection. This drove her to pursue him during World War III in Russia, though he rebuffed her aid to prioritize global stakes. Her persistence entangled her deeper with Academy City’s shadows, including wielding the Anti-Art Attachment, a repurposed nonlethal weapon, despite incurring Aleister Crowley’s curse. Spin-offs expanded her role beyond science-centric conflicts. She unwittingly became a political pawn during Tokiwadai’s student council elections, exposing her naivety toward social schemes. At the Daihaseisai festival, she clashed with forced Level 6 experiments, briefly ascending to a near-angelic state before allies stabilized her. These events crystallized her awareness of systemic corruption and resolve to oppose it. Balancing a tomboyish streak with Tokiwadai’s polished "Ace" persona, she adores Gekota merchandise and whimsical pajamas yet champions justice fiercely. Vulnerabilities surface in insecurities over her physique and frustration at magic’s complexities, grounding her humanity despite her power. Her journey shifts from solitary action to collaborative strategy, embracing aid from Shirai Kuroko, Uiharu Kazari, and Saten Ruiko. Later arcs see her pursuing knowledge beyond Academy City, maturing from local hero to global participant. This trajectory positions her as a nexus between science and magic narratives, though her core remains shielding those she cherishes.