"Di Gi Charat Summer Vacation Special" is a short anime special that centers on the comedic and whimsical adventures of Dejiko, a cat-eared alien girl, and her companions, Puchiko and Gema, as they navigate a summer vacation. The story begins with the trio working at a maid café in Akihabara, where their quirky personalities and antics often lead to humorous situations. Dejiko, the self-proclaimed princess from the planet Di Gi Charat, dreams of becoming a top idol, while Puchiko, her younger companion, is more laid-back and often serves as a foil to Dejiko's energetic nature. Gema, a small, round creature, provides additional comic relief with his mischievous behavior.

The special takes place during a summer break, as the group decides to take a trip to the beach. Their journey is filled with absurd and lighthearted moments, including misunderstandings, playful rivalries, and surreal encounters. Along the way, they cross paths with Rabi~en~Rose, a rival character who often challenges Dejiko, adding a layer of friendly competition to the narrative. The beach setting serves as a backdrop for slapstick humor and exaggerated scenarios, such as Dejiko's attempts to impress others with her idol aspirations, which frequently backfire.

The plot is episodic and driven by character interactions rather than a complex storyline, focusing on the comedic dynamics between the main cast. Dejiko's over-the-top personality and Puchiko's deadpan reactions create a contrast that fuels much of the humor. The special maintains a light, carefree tone, with no significant stakes or dramatic developments, making it a straightforward and entertaining watch. The characters' exaggerated traits and the absurdity of their situations are central to the appeal, offering a playful and nostalgic experience for fans of the "Di Gi Charat" franchise.