"Daimidaler: Prince vs. Penguin Empire" is a 12-episode anime series that blends science fiction, comedy, and ecchi elements. The story centers on Kōichi Madanbashi, a high school student who discovers he possesses a unique ability to generate Hi-ERo particles, a powerful energy source. These particles are harnessed by the Daimidaler, a giant robot designed to combat the Penguin Empire, an alien force threatening Earth. The Penguin Empire, led by the enigmatic Shōma, seeks to conquer humanity by exploiting their perverse desires, which they believe make humans weak.

Kōichi is recruited by Sonan Kōryū, a mysterious woman working for the anti-Penguin Empire organization, Jinrai. To maximize the Daimidaler's power, Kōichi must engage in ecchi activities, such as groping or being groped, to generate Hi-ERo particles. This premise creates a comedic and often absurd tone, as Kōichi struggles with his moral reservations while being thrust into increasingly ridiculous situations. The series frequently parodies mecha anime tropes, with over-the-top battles and exaggerated character reactions.

Supporting characters include Kiriko Kōzuki, Kōichi's childhood friend and love interest, who is unaware of his involvement with Jinrai, and Kyōko Katsuragi, a Jinrai operative who assists Kōichi in battle. The Penguin Empire's forces are composed of anthropomorphic penguins with bizarre personalities and abilities, often serving as comedic foils to the protagonists. As the series progresses, Kōichi and his allies face escalating threats from the Penguin Empire, culminating in a climactic showdown that tests their resolve and teamwork.

The narrative balances its comedic and ecchi elements with occasional moments of character development, particularly regarding Kōichi's growth as a reluctant hero. The series does not shy away from its absurd premise, embracing its over-the-top nature while delivering a straightforward, action-packed story with a mix of humor and fan service.